Quercy Phosphorites Formation

Quercy Phosphorites Formation
Stratigraphic range: latest Bartonian-Late Oligocene
~
Type Formation
Overlies Fissure fillings in karstified Jurassic and Triassic rocks
Lithology
Primary Phosphorite
Location
Coordinates 44.3°N 1.6°E / 44.3; 1.6
Approximate paleocoordinates 42.8°N 2.0°W / 42.8; -2.0
Region Occitanie
Country France
Type section
Named for Quercy
Named by Thévenin
Year defined 1903
Quercy Phosphorites Formation is located in France
Quercy Phosphorites Formation
Quercy Phosphorites Formation (France)

The Quercy Phosphorites Formation (French: Phosphorites du Quercy) is a geologic formation and lagerstätte in Occitanie, southern France. It preserves fossils dated to the Paleogene period (latest Bartonian to Late Oligocene), or MP16 to MP28 zones of the European land mammal age classification, ranging from approximately 38 to 25 Ma.

It qualifies as a Lagerstätte because beside a large variety of mammals, birds, turtles, crocodiles, flora and insects, it also preserves the soft tissues of amphibians and squamates, in addition to their articulated skeleton in what has been called natural mummies.

The genera Quercylurus, Quercymegapodius, Quercypsitta, Quercypodargus, Quercycerta and Quercygama, and species Mosaicomeryx quercyi, Robiacina quercyi, Palaeophyllophora quercyi, Archaeomys quercyi, Eomys quercyi, Eucricetodon quercyi and Tarnomys quercynus, as well as the lizards Paraplacosauriops quercyi and Pseudolacerta quercyini and the insect Palaeortona quercyensis were named after the formation.

Description

The first phosphate deposits in Quercy were discovered in 1869 and published by Daubré and Trutat independently in 1871. The first fossils from the formation were described by Delfortie (1872) and Gervais in the same year and extensively studied by Filhol from 1877 onwards. The first geologic investigation of the formation was performed by Thévenin in 1903, and apart from a description by Gèze in 1938, the paleontological richness was not studied until a team of researchers of the Universities of Montpellier and Paris visited the site in 1965.

The karstified phosphate deposits are found from the Lot and Célé river valleys in the north to the left bank of the Aveyron in the south and from the Villefranche Fault in the east to the lacustrine deposits of the Aquitaine Basin in the west. The formation is found in fissures (karst) incising Jurassic and Triassic rocks east of Cahors. The age of the fossiliferous unit, in which almost 12,000 specimens were found ranges from the MP16 to MP28 zones of the European land mammal age classification. These ages correspond to the latest Bartonian to Chattian, from about 38 to 25 Ma.

Paleontological significance

Paleogeography of the Late Eocene to Oligocene with biodiversity of the Quercy Phosphorites
Exceptional preservation of nerves, digestive tract and stomachal content in the frog-eating salamander Phosphotriton sigei

The Quercy Phosphorites Formation is a highly fossiliferous unit designated as a Lagerstätte due to the excellent preservation of fossils. The phosphorite conserves up to the nerves, digestive tract and stomach content, insect larvae and other elements of the paleobiology in the formation. Nearly all Quercy fly pupae were preserved as isolated endocasts, of which many were still covered by the puparium, the hardened skin of the last larval instar. The formation also straddles the Grande Coupure and shows diversity changes (number of species) of frog, salamander, lizard and snake fossil records across the formation. It is assumed that the Quercy arthropods fossilized by a rapid fixation by phosphate-rich water followed by encrustation and mineralization.

Fossil content

The following fossils have been reported from the formation:

Mammals

Primates
Genus Species Location Stratigraphic position Material Images
Cryptadapis C. tertius An adapiform primate.
Apatotheria
Genus Species Location Stratigraphic position Material Images
Chardinyus C. sp.
Heterohyus H. (Chardinyus) nanus
H. (Gervaisyus) pygmaeus
Gervaisyus G. sp.
Artiodactyls
Genus Species Location Stratigraphic position Material Images
Bachitherium B. guirounetensis
B. lavocati
Cryptomeryx C. gaudryi
Dichobune D. sigei
Dichodon D. vidalenci
Iberomeryx I. matsoui
Mosaicomeryx M. quercyi
Paroxacron P. bergeri
Plesiomeryx P. cadurcensis
'Prodremotherium P. elongatum
Pseudamphimeryx P. salesmei
Robiacina R. lavergnensis
R. quercyi
Tapirulus Tapirulus perrierensis
Carnivora
Genus Species Location Stratigraphic position Material Images
Amphicynodon A. typicus
Cephalogale C. sp.
Cynodyctis C. lacustris neboulensis
Dinailurictis D. bonali
Eofelis E. edwardsii
E. giganteus
cf. E. sp.
Eusmilus E. bidentatus
Mustelictis M. cf. major
M. aff. olivieri
Nimravus N. intermedius
Pachycynodon P. amphictina
P. crassirostris
P. cf. dubius
P. cf. filholi
Palaeogale P. sectoria
Peignictis P. pseudamphictis
Quercylurus Q. major
Wangictis W. tedfordi
Chiroptera
Genus Species Location Stratigraphic position Material Images
Hipposideros Pseudorhinolophus H. P. sehlosseri
H. P. trassounius
H. P. zbrjdi
Leuconoe L. lavocati
Palaeophyllophora P. oltina
P. quercyi
Stehlinia S. bonisi
S. minor
Vaylatsia V. garouillasensis
V. gerscheli
V. gracilis
V. lapradensis
Creodonta
Genus Species Location Stratigraphic position Material Images
Paracynohyaenodon P. magnus
Parapterodon P. lostangensis
Euarchonta
Genus Species Location Stratigraphic position Material Images
Darbonetus D. aubrelongensis
Hyaenodonta
Genus Species Location Stratigraphic position Material Images
Paroxyaena P. pavlovi
Lipotyphla
Genus Species Location Stratigraphic position Material Images
Amphidozotherium A. cayluxi
Darbonetus D. aubrelongensis
D. tuberi
Myxomygale M. antiqua
Saturninia S. beata
S. pelissiei
Marsupials
Genus Species Location Stratigraphic position Material Images
Amphiperatherium A. bourdellense
A. lamandini
Peratherium P. bretouense
P. cayluxi
P. lavergnense
P. perrierense
Perissodactyls
Genus Species Location Stratigraphic position Material Images
Pachynolophus Pachynolophus bretovense
Proeutheria
Genus Species Location Stratigraphic position Material Images
Pseudorhyncocyon Pseudorhyncocyon cayluxi
Rodents
Genus Species Location Stratigraphic position Material Images
Archaeomys A. intermedius
A. quercyi
Bernardia B. marandati
Blainvillimys B. gemellus
B. gousnatensis
B. langei
B. rotundidens
Elfomys E. medius
Eomys Eomys gigas
E. minus
E. quercyi
Eucricetodon E. atavus
E. quercyi
Gliravus G. garouillensis
G. itardiensis
Issiodoromys I. limognensis
I. pauffiensis
Palaeosciurus P. goti
Paradelomys Paradelomys spelaeus
Patriotheridcmys P. altus
P. altus neboulensis
P. sudrei
Pseudoltinomys P. gaillardi
P. major
P. phosphoricus
Sciuromys S. cayluxi
S. rigali
Tarnomys T. quercynus
Theridomys T. ludensis

Birds

Genus Species Location Stratigraphic position Material Images
Ameghinornis A. sp.
Horusornis H. vianeyliaudae
Idiornis I. itardiensis
Itardiornis I. hessae
Leptoganga L. sp.
Necrobyas N. minimus
Nocturnavis N. sp.
Palaeoglaux P. perrierensis
Palaeotodus P. escampsiensis
P. itardiensis
Paleseyvus P. escampensis
Primocolius P. sigei
P. minor
Quercymegapodius Q. brodkorbi
Quercypsitta Q. ivani
Q. sudrei
Recurvirostra R. sanctaeneboulae
Sylphornis S. bretouensis
Ventivorus V. ragei
Caprimulgiformes
Genus Species Location Stratigraphic position Material Images
Euronyctibius E. kurochkini
Quercypodargus Q. olsoni

Reptiles

Crocodiles
  • Alligator gaudryi
Lizards
Snakes
Turtles

Amphibians

Frogs
Salamanders

Insects

Flora