Ancient and Primeval Beech Forests of the Carpathians and Other Regions of Europe

Ancient and Primeval Beech Forests of the Carpathians and Other Regions of Europe
UNESCO World Heritage Site
Beușnița Waterfall in Nera Gorge-Beușnița National Park
Location (see map)
Reference 1133quater
Inscription 2007 (31st Session)
Extensions 2011, 2017, 2021, 2023

Ancient and Primeval Beech Forests of the Carpathians and Other Regions of Europe is a transnational serial nature UNESCO World Heritage Site, encompassing 93 component parts (forests of European beech, Fagus sylvatica) in 18 European countries. Together, the sites protect the largest and least disturbed forests dominated by the beech tree. In many of these stands (especially those in the Carpathians), these forests here were allowed to proceed without interruption or interference since the last ice age. These sites document the undisturbed postglacial repopulation of the species.

Carpathian region

The Primeval Beech Forests of the Carpathians include ten separate massifs located along the 185 km (115 mi) long axis from the Rakhiv mountains and Chornohora ridge in Ukraine over the Poloniny Ridge (Slovakia) to the Vihorlat Mountains in Slovakia. The Ancient Beech Forests of Germany include five locations, cover 4,391 hectares and were added in 2011.

The Carpathian site covers a total area of 77,971.6 ha (192,672 acres), out of which only 29,278.9 ha (72,350 acres) are part of the actual preserved area, while the rest is considered a "buffer zone". Primeval Beech Forests of the Carpathians cover areas of Zakarpattia and Prešov Regions. Over 70% of the site is located in Ukraine. The area includes two national parks, and some habitat controlled areas, mostly in Slovakia. Both national parks, along with a neighboring area in Poland, compose a separate biosphere reserve, the East Carpathian Biosphere Reserve. Besides Havešová, Rožok, and Stužica (all of them located in Bukovské vrchy), there is a fourth component situated in Slovakia, named Kyjovský prales of Vihorlat. Ukrainian locations include Chornohora, Kuziy-Trybushany, Maramarosh, Stuzhytsia–Uzhok, Svydovets, and Uholka–Shyrikyi Luh. Only a few of the ten components are accessible to visitors. Stužica is the only one of three locations in Bukovské vrchy (Slovakia) with available hiking trails. In 2017, UNESCO extended the site, adding forests in Albania, Austria, Belgium, Bulgaria, Croatia, Italy, Romania, Slovenia, and Spain. In 2021 UNESCO extended the site again adding forests in Bosnia and Herzegovina, Czechia, France, Italy, North Macedonia, Poland, Slovakia and Switzerland.

The last intact virgin forest in the temperate latitudes of Europe is to be found in the Carpathians. Trees can live to a hundred years old in these forests, providing an important habitat for organisms such as mushrooms, moss, lichen, insects, rare birds (e.g. capercaillie and black grouse) and mammals (e.g. bats, brown bear, wolf and lynx). Large parts of the forest in the Romanian part of the Carpathians have been lost due to deforestation. The pressure on timber as a resource may increase due to international demand and European companies may start large-scale felling in neighbouring Ukraine. Currently unprotected areas of virgin forest can be permanently preserved in the Ukrainian Carpathians by expanding and reinforcing conservation areas. In the Ukrainian Carpathians there are nine national parks and two biosphere reserves. There is a general ban on tree felling in coniferous forest areas above 1,100 metres. If park administrations are shown to work, management of larger, previously unprotected areas of virgin forest to preserve them on a permanent basis, may occur. There are roughly 100,000 additional hectares of forest which could be integrated into the existing conservation areas.

List of component parts

Map of the distribution of Fagus sylvatica in Europe (green)

The component parts included in the site as of December 2023 are:

Country Serial ID No.
or ID
Component part Component cluster Property [ha] Buffer
zone [ha]
Year
inscrib.
Centre point
coordinates
Carpathian Beech Forest Region
Ukraine 1133quinquies-001
UA_CBR
Chornohora 2,476.80 12,925.00 2007 48°8′25″N 24°23′35″E
1133quinquies-002
UA_CBR
Kuziy-Trybushany 1,369.60 3,163.40 2007 47°56′21″N 24°8′26″E
1133quinquies-003
UA_CBR
Maramarosh 2,243.60 6,230.40 2007 47°56′12″N 24°19′35″E
1133quinquies-004
UA_UNNP
Stuzhytsia-Uzhok 2,532.00 3,615.00 2007 49°4′58″N 22°35′45″E
1133quinquies-005
UA_CBR
Svydovets 3,030.50 5,639.50 2007 48°11′21″N 24°13′37″E
1133quinquies-006
UA_CBR
Uholka-Shyrokyi Luh 11,860.00 3,301.00 2007 48°18′22″N 23°41′46″E
1133quinquies-063
UA_GORG
Gorgany 753.48 4,637.59 2017 48°28′19″N 24°17′58″E
1133quinquies-066
UA_SYNE
Darvaika Synevyr 1,588.46 312.32 2017 48°29′14″N 23°44′56″E
1133quinquies-067
UA_SYNE
Kvasovets 561.62 333.63 2017 48°23′6″N 23°42′46″E
1133quinquies-068
UA_SYNE
Strymba 260.65 191.14 2017 48°27′11″N 23°47′48″E
1133quinquies-069
UA_SYNE
Vilshany 454.31 253.85 2017 48°21′20″N 23°39′36″E
1133quinquies-070
UA_ZACH
Irshavka Zacharovanyi Krai 93.97 1,275.44 2017 48°27′9″N 23°5′23″E
1133quinquies-071
UA_ZACH
Velykyi Dil 1,164.16 2017 48°25′21″N 23°9′42″E
Slovakia 1133quinquies-089
SK-POLO-01
Havešová Poloniny 167.88 6,474.84 2007
2021*
49°0′35″N 22°20′14″E
1133quinquies-090
SK-POLO-02
Rožok 74.37 1,138.89 2007
2021*
48°58′40″N 22°27′52″E
1133quinquies-091
SK-POLO-03
StužicaBukovské Vrchy 1,742.47 5,694.84 2007
2021*
49°5′3″N 22°29′42″E
1133quinquies-092
SK-POLO-04
Udava 455.82 814.69 2007
2021*
49°10′31″N 22°13′39″E
1133quinquies-093
SK-VIHO-02
Vihorlat 1,553.06 854.08 2007
2021*
48°54′56″N 22°11′13″E
Romania 1133quinquies-043
RO_NERE
Cheile Nerei-Beușnița 4,292.27 5,959.87 2017 44°54′19″N 21°48′40″E
1133quinquies-044
RO_SINC
Sinca secular forest (Codrul secular Șinca) 338.24 445.76 2017 45°40′0″N 25°10′14″E
1133quinquies-045
RO_SLAT
Slătioara secular forest (Codrul secular Slătioara) 609.12 429.43 2017 47°26′36″N 25°37′39″E
1133quinquies-046
RO_COZI
Cozia Massif Cozia 2,285.86 2,408.83 2017 45°19′54″N 24°19′32″E
1133quinquies-047
RO_COZI
Lotrisor 1,103.30 2017 45°17′43″N 24°15′33″E
1133quinquies-048
RO_DOMO
Domogled-Coronini-Bedina Domogled –
Valea Cernei
5,110.63 51,461.28 2017 44°56′31″N 22°28′7″E
1133quinquies-049
RO_DOMO
Iauna Craiovei 3,517.36 2017 45°6′31″N 22°34′41″E
1133quinquies-050
RO_DOMO
Ciucevele Cernei 1,104.27 2017 45°14′40″N 22°49′23″E
1133quinquies-051
RO_TIBL
Izvorul Șurii Groșii Țibleșului 210.55 563.57 2017 47°32′59″N 24°11′9″E
1133quinquies-052
RO_TIBL
Preluci 135.82 2017 47°32′5″N 24°13′13″E
1133quinquies-053
RO_IZVO
Izvoarele Nerei 4,677.21 2,494.83 2017 45°7′21″N 22°3′59″E
1133ter-046
RO_STRA
Strâmbu Băiuț 598.14 713.09 2017 47°37′33″N 24°4′23″E
Poland 1133quinquies-085
PL-BIES-01
Border Ridge and
Górna Solinka valley
Bieszczady 1,506.05 24,330.52 2021 49°5′58″N 22°33′24″E
1133quinquies-086
PL-BIES-02
Połonina Wetlińska and Smerek 1,178.03 2021 49°10′51″N 22°30′12″E
1133quinquies-087
PL-BIES-03
Terebowiec stream valley 201.00 2021 49°5′37″N 22°43′29″E
1133quinquies-088
PL-BIES-04
Wołosatka stream valley 586.66 2021 49°4′1″N 22°44′41″E
Polonic-Podoloc-Moldovan Beech Forest Region
Ukraine 1133quinquies-064
UA_ROZT
Roztochchya 384.81 598.21 2017 49°57′44″N 23°38′58″E
1133quinquies-065
UA_SATA
Satanіvska Dacha 212.01 559.37 2017 49°10′26″N 26°14′56″E
Baltic Beech Forest Region
Germany 1133quinquies-007
DE_JASM
Jasmund 492.50 2,510.50 2011 54°32′53″N 13°38′43″E
1133quinquies-008
DE_SERR
Serrahn 268.10 2,568.00 2011 53°20′24″N 13°11′52″E
1133quinquies-009
DE_GRUM
Grumsin 590.10 274.30 2011 52°59′11″N 13°53′44″E
Subatlantic-Hercynic Beech Forest Region
Germany 1133quinquies-010
DE_HAIN
Hainich 1,573.40 4,085.40 2011 51°04′43″N 10°26′08″E
1133quinquies-011
DE_KELL
Kellerwald 1,467.10 4,271.40 2011 51°08′43″N 8°58′25″E
Czech Republic 1133quinquies-075
CZ-JIZE-01
Jizera Mountains 444.81 2,279.40 2021 50°51′30″N 15°9′20″E
France 1133quinquies-077
FR-GRAN-01
Grand Ventron 319.00 1,328.00 2021 47°58′20″N 6°56′23″E
Switzerland 1133quinquies-073
CH-BETT-01
Bettlachstock Forest
(Forêt de la Bettlachstock)
195.43 1,094.16 2021 47°13′22″N 7°24′43″E
Alpic Beech Forest Region
Austria 1133quinquies-014
AT_DUER
Dürrenstein-Lassingtal 3,685.27 3,089.78 2017
2023*
47°46′12″N 15°2′51″E
1133quinquies-015
AT_KALK
Hintergebirge Kalkalpen 2,946.20 14,197.24 2017 47°44′58″N 14°28′56″E
1133quinquies-016
AT_KALK
Bodinggraben 890.89 2017 47°47′14″N 14°21′12″E
1133quinquies-017
AT_KALK
Urlach 264.82 2017 47°48′15″N 14°14′22″E
1133quinquies-018
AT_KALK
Wilder Graben 1,149.75 2017 47°50′00″N 14°26′1″E
Switzerland 1133quinquies-074
CH-LODA-01
Forest reserves of Lodano, Busai and Soladino Valleys 806.78 2,330.74 2021 46°15′49″N 8°39′11″E
France 1133quinquies-076
FR-CHAP-01
Chapitre 371.30 41.65 2021 44°38′4″N 5°59′55″E
Atlantic Beech Forest Region
Belgium 1133quinquies-019
BE_SONI
Forest Reserve “Joseph Zwaenepoel” Sonian Forest
(Forêt de Soignes)
187.34 4,650.86 2017 50°45′23″N 4°25′00″E
1133quinquies-020
BE_SONI
Grippensdelle A 24.11 2017 50°46′54″N 4°25′36″E
1133quinquies-021
BE_SONI
Grippensdelle B 37.38 2017 50°47′1″N 4°25′57″E
1133quinquies-022
BE_SONI
Réserve forestière du Ticton A 13.98 2017 50°44′3″N 4°26′13″E
1133quinquies-023
BE_SONI
Réserve forestière du Ticton B 6.50 2017 50°43′37″N 4°25′51″E
Moesian-Balcanic Beech Forest Region
Albania 1133quinquies-012
AL_LUMI
Lumi i gashit 1,261.52 8,977.48 2017 42°25′36″N 20°05′45″E
1133quinquies-013
AL_RRAJ
Rrajca 2,129.45 2,569.75 2017 41°12′11″N 20°30′2″E
North Macedonia 1133quinquies-084
MN-DLAB-01
Dlaboka Reka 193.27 234.70 2021 41°45′47″N 20°35′16″E
Bulgaria 1133quinquies-024
BG_BALK
Boatin Reserve Central Balkan 1,226.88 851.22 2017 42°48′10″N 24°16′9″E
1133quinquies-025
BG_BALK
Tsarichina Reserve 1,485.81 1,945.99 2017 42°46′32″N 24°24′18″E
1133quinquies-026
BG_BALK
Kozya stena Reserve 644,43 289.82 2017 42°47′47″N 24°31′2″E
1133quinquies-027
BG_BALK
Steneto Reserve 2,466.10 1,762.01 2017 42°44′43″N 24°42′26″E
1133quinquies-028
BG_BALK
Stara Reka Reserve 591.20 1,480.04 2017 42°42′11″N 24°49′8″E
1133quinquies-029
BG_BALK
Dzhendema Reserve 1,774.12 2,576.63 2017 42°41′44″N 24°58′23″E
1133quinquies-030
BG_BALK
Severen Dzhendem Reserve 926.37 1,066.47 2017 42°44′44″N 24°56′5″E
1133quinquies-031
BG_BALK
Peeshti skali Reserve 1,049.10 968.14 2017 42°45′54″N 25°4′29″E
1133quinquies-032
BG_BALK
Sokolna Reserve 824.90 780.55 2017 42°41′52″N 25°8′18″E
Illyric Beech Forest Region
Croatia 1133quinquies-033
HR_HAJD
Forest reserves Hajdučki i Rožanski kukovi 1,289.11 9,869.25 2017 44°45′59″N 15°0′39″E
1133quinquies-034
HR_PAKL-01
Paklenica National Park
(formerly
PNP - Suva draga-Klimenta and
PNP - Oglavinovac-Javornik)
2,036.81 824.86 2017
2023*
44°20′26″N 15°30′1″E
Slovenia 1133quinquies-055
SI_KROK
Krokar 74.50 47.90 2017 45°32′31″N 14°46′8″E
1133quinquies-056
SI_SNEZ
Snežnik-Ždrocle 720.24 128.80 2017 45°35′5″N 14°27′19″E
Bosnia and
Herzegovina
1133quinquies-072
BH-JANJ-01
Prašuma Janj 295.04 380.74 2021 44°8′48″N 17°16′52″E
Central Mediterranean Beech Forest Region
Italy 1133quinquies-035
IT_ABRU
Cervara Valley (Valle Cervara) Abruzzo, Lazio & Molise 119.70 751.61 2017 41°49′56″N 13°43′43″E
1133quinquies-036
IT_ABRU
Selva Moricento 192.70 2017 41°50′49″N 13°42′20″E
1133quinquies-037
IT_ABRU
Coppo del Morto 104.71 415.51 2017 41°51′37″N 13°50′48″E
1133quinquies-038
IT_ABRU
Coppo del Principe 194.49 446.62 2017 41°47′15″N 13°44′39″E
1133quinquies-039
IT_ABRU
Fondillo Valley (Val Fondillo) 325.03 700.95 2017 41°45′15″N 13°53′9″E
1133quinquies-081
IT-POLL-01
Cozzo Ferriero Pollino National Park 95.75 2,851.83 2017
2021*
39°54′19″N 16°6′4″E
1133quinquies-082
IT-POLL-02
Pollinello 477.94 2021 39°53′43″N 16°11′54″E
1133quinquies-083
IT-VALL-01
Infernale Valley (Valle Infernale) 320.79 2,191.36 2021 38°7′55″N 15°57′41″E
1133quinquies-079
IT-FUMB-01
Falascone Umbra Forest
(Foresta Umbra)
254.30 3,486.29 2017
2021*
41°48′21″N 15°58′41″E
1133quinquies-080
IT-FUMB-02
Pavari-Sfilzi 667.13 2021 41°50′20″N 16°1′25″E
1133quinquies-040
IT_CIMI
Monte Cimino 57.54 87.96 2017 42°24′31″N 12°12′11″E
1133quinquies-041
IT_RASC
Monte Raschio 73.73 54.75 2017 42°10′25″N 12°9′40″E
1133quinquies-042
IT_SASS
Sasso Fratino 781.43 6,936.64 2017 43°50′40″N 11°48′11″E
Pyrenaic-Iberian Beech Forest Region
Spain 1133quinquies-057
ES_AYLL
Tejera Negra Beech forests of Ayllón
(Hayedos de Ayllón)
255.52 13,880.86 2017 41°14′43″N 3°23′19″W
1133quinquies-058
ES_AYLL
Montejo 71.79 2017 41°6′44″N 3°29′58″W
1133quinquies-059
ES_NAVA
Lizardoia Beech forests of Navarra
(Hayedos de Navarra)
63.97 24,494.52 2017 43°0′23″N 1°6′46″W
1133quinquies-060
ES_NAVA
Aztaparreta 171.06 2017 42°54′39″N 0°48′58″W
1133quinquies-061
ES_PICO
Cuesta Fría Beech forests of
Picos de Europa

(Hayedos de Picos de Europa)
213.65 14,253.00 2017 43°10′21″N 4°59′16″W
1133quinquies-062
ES_PICO
Canal de Asotin 109.58 2017 43°10′16″N 4°53′21″W
France 1133quinquies-078
FR-MASS-01
Massane 239.50 1,432.30 2021 42°28′58″N 3°1′45″E

* Only boundary modification.

Udava (Bukovské Vrchy, Slovakia) was created by modifying the boundary of Stužica before. Falascone (Italy) in turn was named previously Umbra Forest (Foresta Umbra).

This is a transnational serial nature UNESCO World Heritage Site, encompassing 94 component parts (forests) in 18 European countries. The total area of component parts is 98,125.15 ha (242,472.5 acres), including buffer zones 392,846.02 ha (970,743.7 acres). The largest total area of component parts is in Ukraine and Romania – 54% total (41% including buffer zones).

Forms of protection

Component parts of this property overlap more or less with various protected areas like national parks, nature reserves, Natura 2000 network.
Abbreviations:
C – count of component parts in a given cluster
N – overlapping with Natura 2000 areas

Country Component cluster C Protected area N Region
Albania Lumi i Gashit 1 Strict nature reserve of Gashi River Kukës
Rrajca 1 Shebenik – Jabllanicë National Park Elbasan
Austria Dürrenstein-Lassingtal 1 Wilderness Area Dürrenstein Yes Lower Austria
Kalkalpen 4 Kalkalpen National Park Yes Upper Austria
Belgium Sonian Forest
(Forêt de Soignes)
5 3 strict forest reserves within Sonian Forest Protected Landscape – “Joseph Zwaenepoel”, Grippensdelle and Ticton Yes Flanders, Brussels Capital Region, Wallonia
Bosnia and Herzegovina Prašuma Janj 1 Prašuma Janj strict nature reserve Bosanska Krajina
Bulgaria Central Balkan 9 9 strict reserves in the Central Balkan National Park – Boatin, Dzhendema, Kozya stena, Peeshti skali, Severen Dzhendem, Sokolna, Stara reka, Steneto and Tsarichina Yes Lovech, Plovdiv, Stara Zagora, Gabrovo (central Balkan Mountains)
Croatia Hajdučki i Rožanski kukovi 1 Hajdučki and Rožanski Kukovi strict reserve within Northern Velebit National Park Yes Lika-Senj
Paklenica National Park 2 Paklenica National Park Yes Lika-Senj, Zadar
(Dinaric region, or Dinaric Alps)
Czech Republic Jizera Mountains 1 Jizerskohorské bučiny National Nature Reserve Yes Liberec
France Chapitre 1 Chapitre-Petit-Buëch biological strict reserve Yes Provence-Alpes-Côte d'Azur
Grand Ventron 1 Massif du Grand Ventron National Nature Reserve Yes Grand Est
Massane 1 Massane National Nature Reserve Yes Occitanie
Germany Grumsin 1 Schorfheide-Chorin Biosphere Reserve Yes Brandenburg
Hainich 1 Hainich National Park Yes Thuringia
Jasmund 1 Jasmund National Park Yes Mecklenburg-Vorpommern
Kellerwald 1 Kellerwald-Edersee National Park Yes Hesse
Serrahn 1 Müritz National Park Yes Mecklenburg-Vorpommern
Italy Abruzzo, Lazio & Molise 5 Abruzzo, Lazio and Molise National Park Yes Abruzzo
Umbra Forest (Foresta Umbra) 2 Strict reserves within Gargano National Park – Sfilzi, Falascone and Foresta Umbra Yes Apulia
Monte Cimino 1 Regional Natural Monument and municipal strict reserve Yes Lazio
Monte Raschio 1 Regional Natural Park Bracciano-Martignano Yes Lazio
Pollino National Park 2 Cozzo Ferriero and Pollinello strict reserves within Pollino National Park Yes Calabria, Basilicata
Sasso Fratino 1 Sasso Fratino strict reserve in Foreste Casentinesi, Monte Falterona, Campigna National Park Yes Emilia-Romagna, Tuscany
Infernale Valley (Valle Infernale) 1 Valle Infernale strict reserve within Aspromonte National Park Yes South Calabria
North Macedonia Dlaboka Reka 1 Forest reserves within Mavrovo National Park Mavrovo and Rostuša and Gostivar
Poland Bieszczady 4 Bieszczady National Park, East Carpathian Biosphere Reserve Yes Podkarpackie Voivodeship
Romania Cheile Nerei-Beușnița 1 Strictly protected zone of the Cheile Nerei-Beușnița National Park Yes Caraș-Severin
Sinca secular forest
(Codrul secular Șinca)
1 Strictly protected through the forest management plan Yes Brașov
Slătioara secular forest
(Codrul secular Slătioara)
1 Part of Codrul Secular Slătioara forest reserve, Secular Meadows Todirescu floristic reserve Yes Suceava
Cozia 2 Cozia National Park Yes Vâlcea
Domogled – Valea Cernei 3 Domogled-Valea Cernei National Park Yes Mehedinți, Caraș-Severin, Gorj
Groșii Țibleșului 2 Strictly protected through the forest management plan Yes Maramureș
Izvoarele Nerei 1 Izvoarele Nerei strict reserve within Semenic-Cheile Carașului National Park Yes Caraș-Severin
Strâmbu Băiuț 1 Strictly protected through the forest management plan Yes Maramureș
Slovakia Poloniny 4 National nature reserves Stužica, Jarabá skala, Pľaša and Borsukov vrch (Stužica – Bukovské Vrchy), Rožok, Havešová, nature reserve Udava within Poloniny National Park, East Carpathian Biosphere Reserve Yes Prešov
Vihorlat 1 Vihorlat national nature reserve within Vihorlat Protected Landscape Area Yes Prešov
Slovenia Krokar 1 Forest reserve Virgin forest Krokar Yes Kočevje
Snežnik–Ždrocle 1 Snežnik–Ždrocle forest reserve Yes Inner Carniola
Spain Beech forests of Ayllón
(Hayedos de Ayllón)
2 Tejera Negra: Sierra Norte de Guadalajara Natural Park, Montejo: Sierra del Rincón Biosphere Reserve Yes Castilla-La Mancha
Beech forests of Navarra
(Hayedos de Navarra)
2 Integral (strict) reserves of Lizardoia and Aztaparreta Yes Navarre
Beech forests of
Picos de Europa
(Hayedos de Picos de Europa)
2 Picos de Europa National Park, Biosphere Reserve Yes Castile and León
Switzerland Bettlachstock Forest
(Forêt de la Bettlachstock)
1 Forest reserve Bettlachstock–Hasenmatt Canton of Solothurn
Forest reserves of Lodano, Busai and Soladino Valleys 1 Lodano Valley forest reserve, Busai and Soladino Valleys forest reserve Canton of Ticino
Ukraine Chornohora 1 Carpathian Biosphere Reserve Zakarpattia
Kuziy – Trybushany 1 Carpathian Biosphere Reserve Zakarpattia
Maramarosh 1 Carpathian Biosphere Reserve Zakarpattia
Stuzhytsia – Uzhok 1 Uzhanian National Nature Park Zakarpattia
Svydovets 1 Carpathian Biosphere Reserve Zakarpattia
Uholka – Shyrokyi Luh 1 Carpathian Biosphere Reserve Zakarpattia
Synevyr 4 Synevyr National Nature Park Zakarpattia
Zacharovanyi Krai 2 Zacharovany Krai National Nature Park Zakarpattia
Gorgany 1 Gorgany Nature Reserve Ivano-Frankivsk
Roztochchya 1 Roztochchya Nature Reserve, Roztochia Biosphere Reserve Lviv
Satanіvska Dacha 1 Podilski Tovtry National Nature Park Khmelnytskyi

Tentative list

This forest is placed on the World Heritage Tentative list as a proposal for the expansion of the Ancient and Primeval Beech Forests of the Carpathians and Other Regions of Europe site:

Montenegro

Serbia

Gallery